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The only way to get an older OS directly from Apple is to have previously 'purchased it' - even if it was free - then it will appear in your purchase history in the App Store.
If it's not in your purchase history, then you'll have to get it from a trustworthy friend [or less-trustworthy torrent etc] 10.10.
Now that mac OS 10.11 (El Capitan) has been released, what can I do to upgrade to mac OS 10.10.5 if/when I wish to?
When I am in the Apple app store the only option I see mac OS 10.11 as an upgrade option; screenshot below.
You get the latest features and you get the best security.
If you want to upgrade to Mojave you can simply search for Mojave in the Mac App Store and it comes up and you can upgrade from there. As a matter of fact if you don't see it here and you can't get to it in the Mac App Store that probably means your Mac is too old. Well, Apple has you covered because they have links that go directly to these.
Apple has pages for High Sierra, Sierra and El Capitan than include links directly to the hidden Mac App Store pages with those downloads. On this upgrade page here for High Sierra it has the requirements for upgrading to High Sierra. If we go back here we can see there was the link for El Capitan and you can go and click on that here and get to the El Capitan link in the Mac App Store.The logic is, I assume that mac OS 10.10.5 is more stable that mac OS 10.10 when it was first released, so I would like to put a 10.10.5 upgrade on my plate of options if/when I have the time and patience to deal with.I would like to avoid mac OS 10.11 (El Capitan) for a while if possible.Since I had previously downloaded and associated each of these versions with my Apple ID (with the exception of Mojave), all the App Store download links worked for me.The currently existing answers don't answer the questions posed in your last paragraph, so I'll cover that here.